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Abstract Studying the centennial or millennial timescale response of large rivers to changing patterns in precipitation, discharge, flood intensity and recurrence, and associated sediment erosion is critical for understanding long‐term fluvial geomorphic adjustment to climate. Long hydrographs, maintaining reliable Flow Duration Curves (FDCs), are a fundamental input for such simulations; however, recorded discharge series rarely span more than a few decades. The absence of robust methodologies for generating representative long‐term hydrographs, especially those incorporating coarse temporal resolution or lacking continuous simulations, is therefore a fundamental challenge for climate resilience. We present a novel approach for constructing multi‐century hydrographs that successfully conserve the statistical, especially frequency analysis, and stochastic characteristics of observed hydrographs. This approach integrates a powerful combination of a weather generator with a fine disaggregation technique and a continuous rainfall‐runoff transformation model. We tested our approach to generate a statistically representative 300‐year hydrograph on the Ninnescah River Basin in Kansas, using a satellite precipitation data set to address the considerable gaps in the available hourly observed data sets. This approach emphasizes the similarities of FDCs between the observed and generated hydrographs, exhibiting a reasonably acceptable range of average absolute deviation between 6% and 18%. We extended this methodology to create projected high‐resolution hydrographs based on a range of climate change scenarios. The projected outcomes present pronounced increases in the FDCs compared to the current condition, especially for more distant futures, which necessitates more efficient adaptation strategies. This approach represents a paradigm shift in long‐term hydrologic modeling. 

Summary Joshua trees are long‐lived perennial monocots native to the Mojave Desert in North America. Composed of two species,Yucca brevifoliaandY. jaegeriana(Asparagaceae), Joshua trees are imperiled by climate change, with decreases in suitable habitat predicted under future climate change scenarios. Relatively little is understood about the ecophysiology of Joshua trees across their range, including the extent to which populations are locally adapted or phenotypically plastic to environmental stress.Plants in our common gardens showed evidence of Crassulacean acid metabolism photosynthesis (CAM) in a pilot experiment, despite no prior report of this photosynthetic pathway in these species. We further studied the variation and strength of CAM within a single common garden, measuring seedlings representing populations across the range of the two species.A combination of physiology and transcriptomic data showed low levels of CAM that varied across populations but were unrelated to home environmental conditions. Gene expression confirmed CAM activity and further suggested differences in carbon and nitrogen metabolism betweenY. brevifoliaandY. jaegeriana.Together the results suggest greater physiological diversity between these species than initially expected, particularly at the seedling stage, with implications for future survival of Joshua trees under a warming climate.